:root {
  --ink: #1f2a21;
  --muted: #5e6b61;
  --accent: #4ca86c;
  --accent-dark: #22613b;
  --paper: rgba(255, 252, 245, 0.94);
  --paper-strong: rgba(255, 252, 245, 0.98);
  --line: rgba(31, 42, 33, 0.14);
  --shadow: 0 18px 60px rgba(0,0,0,0.30);
}

* {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
  width: 100%;
  overflow-x: hidden;
}

body {
  width: 100%;
  min-width: 0;
  margin: 0;
  overflow-x: hidden;
  font-family: Georgia, "Times New Roman", serif;
  color: var(--ink);
  background: #07111d;
  line-height: 1.6;
}

/* Site-wide histology background */
body::before {
  content: "";
  position: fixed;
  inset: 0;
  background:
    linear-gradient(180deg, rgba(2, 10, 18, 0.46) 0%, rgba(2, 10, 18, 0.64) 100%),
    url("../img/area-x-histology-background.jpg") center center / cover no-repeat;
  z-index: -2;
}

body::after {
  content: "";
  position: fixed;
  inset: 0;
  background:
    radial-gradient(circle at 50% 0%, rgba(76,168,108,0.22), rgba(0,0,0,0) 42%),
    linear-gradient(180deg, rgba(0,0,0,0.12), rgba(0,0,0,0.34));
  z-index: -1;
}

a {
  color: var(--accent-dark);
  text-decoration-thickness: 1px;
  text-underline-offset: 3px;
}

a:hover {
  color: var(--accent);
}

.wrap {
  width: calc(100% - 48px);
  max-width: 1000px;
  margin-left: auto;
  margin-right: auto;
}

.site-nav {
  padding: 22px 0 16px;
  background: rgba(255, 252, 245, 0.90);
  border-bottom: 1px solid rgba(255,255,255,0.36);
  backdrop-filter: blur(8px);
}

.nav-inner {
  display: flex;
  justify-content: space-between;
  align-items: center;
  gap: 18px;
  flex-wrap: wrap;
}

.brand {
  font-size: 1.05rem;
  letter-spacing: 0.02em;
  text-decoration: none;
  color: var(--ink);
  font-weight: 700;
}

.nav-links {
  display: flex;
  gap: 22px;
  align-items: center;
  flex-wrap: wrap;
  font-size: 0.98rem;
}

.nav-links a {
  color: var(--ink);
  text-decoration: none;
  border-bottom: 1px solid transparent;
}

.nav-links a:hover {
  border-color: var(--accent);
}

.hero {
  padding: 44px 0 4px;
}

main {
  position: relative;
  z-index: 2;
  margin-top: 0;
}

.section {
  margin: 28px auto;
  padding: clamp(23px, 4vw, 34px);
  background: var(--paper);
  border: 1px solid rgba(255,255,255,0.50);
  border-radius: 22px;
  box-shadow: 0 12px 36px rgba(0,0,0,0.24);
  backdrop-filter: blur(7px);
}

.hero-card {
  width: 100%;
  margin: 0 auto;
  background: rgba(255, 252, 245, 0.92);
  border: 1px solid rgba(255,255,255,0.64);
  box-shadow: var(--shadow);
}

.research-figure {
  grid-column: 1 / -1;
  margin: 28px 0 0;
  width: 100%;
  overflow: visible;
}

.research-figure img {
  display: block;
  width: 100%;
  max-width: none;
  height: auto;
  margin-left: 0%;
  border-radius: 16px;
}

.intro-grid,
.about-grid {
  display: grid;
  grid-template-columns: minmax(0, 1fr) 265px;
  gap: 30px;
  align-items: start;
}

.about-grid {
  grid-template-columns: 265px minmax(0, 1fr);
}

.media-photo {
  margin: 0;
}

.media-photo img {
  display: block;
  width: 240px;
  max-width: 100%;
  aspect-ratio: 1 / 1;
  object-fit: cover;
  border-radius: 24px;
  border: 1px solid var(--line);
  box-shadow: 0 14px 34px rgba(0,0,0,0.20);
}

.headshot img {
  object-position: 50% 24%;
}

.finch-photo img {
  object-position: 52% 44%;
}

.caption {
  margin: 10px 0 0;
  color: var(--muted);
  font-size: 0.86rem;
  line-height: 1.35;
}

.finch-photo {
  width: 240px;
  max-width: 100%;
}

.finch-photo .caption {
  width: 100%;
  margin: 10px auto 0;
  text-align: center;
}

.kicker {
  margin: 0 0 12px;
  color: var(--accent-dark);
  text-transform: uppercase;
  letter-spacing: 0.14em;
  font-size: 0.78rem;
  font-weight: 700;
  font-family: system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
}

h1 {
  max-width: 100%;
  margin: 0;
  font-size: clamp(2.3rem, 6.1vw, 3.85rem);
  line-height: 1.03;
  letter-spacing: -0.035em;
  text-wrap: balance;
}

.tagline {
  margin: 22px 0 0;
  max-width: 560px;
  font-size: clamp(1.08rem, 2vw, 1.28rem);
  line-height: 1.42;
}

.quick-links {
  display: flex;
  gap: 10px;
  flex-wrap: wrap;
  margin-top: 26px;
}

.button {
  display: inline-block;
  max-width: 100%;
  padding: 9px 13px;
  border: 1px solid rgba(34, 97, 59, 0.34);
  border-radius: 999px;
  background: rgba(255,255,255,0.72);
  color: var(--accent-dark);
  text-decoration: none;
  font-family: system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  font-size: 0.9rem;
  font-weight: 650;
}

.button:hover {
  border-color: var(--accent);
  background: white;
}

h2 {
  margin: 0 0 14px;
  font-size: 1.42rem;
  line-height: 1.2;
}

h3 {
  margin: 0 0 6px;
  font-size: 1.05rem;
}

p {
  margin: 0 0 16px;
}

p:last-child {
  margin-bottom: 0;
}

.section,
.hero-card,
.card,
.pub-list li,
.button {
  overflow-wrap: anywhere;
}

.cards {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 12px;
}

.card {
  padding: 16px;
  border: 1px solid var(--line);
  border-radius: 16px;
  background: rgba(255,255,255,0.55);
}

.card p {
  color: var(--muted);
  font-size: 0.95rem;
}

.news p,
.pub-list li {
  margin-bottom: 11px;
}

.news h3 {
  margin: 20px 0 8px;
  font-size: 1.05rem;
  letter-spacing: 0.02em;
  color: var(--text);
}

.featured-news {
  padding: 14px 16px;
  border: 1px solid var(--line);
  border-radius: 16px;
  background: rgba(255, 255, 255, 0.08);
}

.featured-news p {
  margin-bottom: 0;
  font-size: 1.02rem;
}

.news-divider {
  border: 0;
  border-top: 1px solid var(--line);
  margin: 24px 0 18px;
}


.pub-list {
  margin: 0;
  padding-left: 21px;
}

.page-title {
  font-size: clamp(2.1rem, 6.2vw, 3.25rem);
  margin: 0 0 14px;
  letter-spacing: -0.03em;
}

.lead {
  font-size: 1.08rem;
  color: var(--ink);
  max-width: 650px;
}

.small-note {
  color: var(--muted);
  font-size: 0.93rem;
}

.publications-page {
  margin-top: 28px;
  background: var(--paper-strong);
}

.full-pub-list li {
  padding-bottom: 15px;
  margin-bottom: 15px;
  border-bottom: 1px solid var(--line);
}

.full-pub-list li:last-child {
  border-bottom: 0;
  padding-bottom: 0;
}

.pub-citation {
  margin-bottom: 8px;
}

.pub-links {
  display: flex;
  gap: 8px;
  flex-wrap: wrap;
  font-family: system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  font-size: 0.84rem;
}

.pub-links a {
  display: inline-block;
  padding: 4px 9px;
  border: 1px solid rgba(34, 97, 59, 0.22);
  border-radius: 999px;
  background: rgba(255,255,255,0.62);
  color: var(--accent-dark);
  text-decoration: none;
  font-weight: 650;
}

.pub-links a:hover {
  border-color: var(--accent);
  background: white;
}


.footer {
  padding: 26px 0 40px;
  color: rgba(255,255,255,0.82);
  font-size: 0.92rem;
  text-shadow: 0 1px 2px rgba(0,0,0,0.5);
}

.footer-links {
  display: flex;
  gap: 16px;
  flex-wrap: wrap;
  margin-bottom: 10px;
  font-family: system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
}

.footer-links a {
  color: rgba(255,255,255,0.92);
  text-decoration: none;
}

.footer-links a:hover {
  color: #fff;
  text-decoration: underline;
}

@media (max-width: 840px) {
  body::before {
    background:
      linear-gradient(180deg, rgba(2, 10, 18, 0.50) 0%, rgba(2, 10, 18, 0.70) 100%),
      url("../img/area-x-histology-background.jpg") center center / cover no-repeat;
  }

  .wrap {
    max-width: 760px;
  }

  .intro-grid,
  .about-grid,
  .cards {
    grid-template-columns: 1fr;
  }

  .media-photo img {
    width: 240px;
  }
}

@media (max-width: 520px) {
  .wrap {
    width: calc(100% - 28px);
    max-width: none;
  }

  .site-nav {
    padding-top: 18px;
  }

  .nav-inner {
    align-items: flex-start;
  }

  .hero {
    padding-top: 28px;
  }

  .hero-card,
  .section {
    border-radius: 18px;
    padding: 22px 18px;
  }

  .kicker {
    font-size: 0.7rem;
    letter-spacing: 0.10em;
  }

  h1 {
    font-size: clamp(2.15rem, 12.4vw, 3.0rem);
    letter-spacing: -0.025em;
  }

  .tagline {
    font-size: 1.05rem;
  }

  .quick-links {
    gap: 8px;
  }

  .button {
    font-size: 0.86rem;
    padding: 8px 11px;
  }

  .media-photo img {
    width: 220px;
  }
}